NEXUS
[File created by TreeBASE:  7/26/07  21:08:41]
[Matrix accession#: M2332]

BEGIN TAXA;
	TITLE M2332;
	DIMENSIONS  NTAX=13;
	TAXLABELS
		Corytophanes_hernandezi	Corytophanes_percarinatus	Corytophanes_cristatus	Laemanctus_serratus	Laemanctus_longipes	Basiliscus_plumifrons	Basiliscus_galeritus	Basiliscus_vittatus	Basiliscus_basiliscus	Polychrus_acutirostris	Tropidurus_etheridgei	Sauromalus_obesus	Elgaria_panamintina
	;
END;

BEGIN CHARACTERS;
	TITLE M2332;
	DIMENSIONS NCHAR=89;
	FORMAT SYMBOLS = " 0 1 2 3 " MISSING = ? GAP = - ;
	CHARSTATELABELS
		1 Lacrimal_5 / present absent, 2 Skull_rugosity_7 / absent_or_restricted_to_frontal_bone extensive, 3 'Jugal, squamosal contact 8' / not_or_barely_in_contact broadly_juxtaposed, 4 Postfrontal_9 / present extremely_small_or_absent, 5 Parietal_roof_shape_10 / trapezoidal 'V or Y-shaped' 'Y-shaped with posteriorly directed median crest developed postembryonically' 'Y-shaped with median crest developed embrionically', 6 Parietal_foramen_11 / at_frontoparietal_suture_or_in_parietal entirely_within_the_frontal, 7 Supratemporal_12 / mostly_on_the_lateral_or_ventral_surface_of_the_supratemporal_process_of_the_parietal mostly_on_the_medial_surface_of_the_supratemporal_process_of_the_parietal mostly_in_a_groove_in_the_ventral_margin_supratemporal_process_of_the_parietal, 8 Osseous_labyrinth_13 / low_to_moderate_elevation_of_the_osseous_labyrinth_above_the_general_level_of_the_opisthotics high_elevation_above_the_general_level_of_opisthotics, 9 Endolymphatic_sacs_14 / do_not_extend_outside_of_otic_capsule_into_nuchal_musculature extend_in_nuchal_musculature, 10 'Splenial, anterior extent 21' / extends_anteriorly_to_or_beyond_0.5_length_of_tooth_row does_not_extend_anteriorly_more_than_0.5_length_of_tooth_row does_not_extend_anteriorly_more_than_0.17_length_of_tooth_row, 11 'Splenial, posterior extent 22' / terminates_posteriorly_anterior_to_anterior_edge_of_mandibular_fossa 'terminates posterior to, or at anterior edge of mandibular fossa', 12 'Angular, condition of contact with splenial 23' / 'angular large; suture with splenial on lingual face' 'angular small; suture with splenial on ventral or labial face', 13 Posterior_mylohyoid_foramen_24 / anterior_to_or_approximately_at_the_level_of_superior_apex_of_coronoid between_level_of_superior_apex_of_coronoid_and_anterior_end_of_adductor_fossa posterior_to_anterior_end_of_adductor_fossa, 14 Ceratobranchials_29 / second_not_reaching_clavicles reaching, 15 Clavicle_30 / 'flat, with wide lateral flange' flange_small_or_absent, 16 Insertion_of_clavicle_31 / on_suprascapula on_scapula, 17 Interclavicle_32 / anterior_process_absent anterior_process_well_developed, 18 'Sternum, anterior extent 33' / sternum_does_not_approach_junction_of_posterior_and_lateral_processes_of_interclavicle_closely approaches, 19 Caudal_vertebral_type_34 / Sceloporus_condition Iguana_condition Basiliscus_condition anole_condition, 20 Scapular_fenestra_35 / present absent, 21 Posterior_coracoid_fenestra_36 / present absent, 22 Median_enlarged_sternal_fontanelles_37 / 'absent or small, often hiden by interclavicle' 'present, large, and not paired' 'present, large, and paired', 23 Cervical_ribs_38 / first_pair_on_vertebra_number_3 on_number_4 on_number_5, 24 Number_of_sternal_ribs_39 / 4. 3. 2_or_fewer, 25 Postxiphisternal_inscriptional_ribs_40 / all_attached_proximally_to_dorsal_ribs_and_none_are_confluent_midventrally one_or_more_pairs_attached_to_dorsal_ribs_and_are_confluent_midventrally none_attached_to_dorsal_ribs_or_continuous_midventrally, 26 Tail_autotomy_fracture_planes_41 / present absent, 27 Interparietal_scale_42 / small_or_absent 'large; as wide as interorbital space', 28 Subocular_scale_45 / at_least_one_scale_below_the_eye_conspicuously_enlarged scales_below_the_eye_subequal, 29 'Mid-dorsal scale row 46' / present absent, 30 Gular_fold_47 / complete_medially incomplete_or_absent, 31 Femoral_pores_48 / present absent, 32 'Nasal chamber, S-condition 54' / primitive_condition 'S-condition, septomaxilla plow-share shaped', 33 'Nasal chamber, fusion of nasal concha to roof of nasal chamber 55' / primitive_condition fusion_of_concha_to_roof_of_nasal_chamber, 34 'Nasal chamber, anole condition 56' / primitive_condition nasal_concha_lost, 35 Dorsal_shank_muscle_innervation_59 / 'A-condition (peroneus)' 'B-condition (interosseus)', 36 'Hemipenis, capitation and sulci 61' / unicapitate_or_weakly_bilobate_without_distinctly_divided_sulci bilobate_with_distinctly_divided_sulci strongly_bicapitate, 37 'Hemipenis, m. retractor lateralis posterior 62' / not_substantially_situated_within_the_hemipenial_sheath substantially, 38 'Hemipenis, dorsal accessory sheath 64' / absent present, 39 Colic_septa_65 / absent present, 40 Roof_of_nasal_chamber_1 / nasal_capsule_entirely_covered_by_frontal_and_nasals nasal_capsule_uncovered_at_junction_of_frontal_and_nasals, 41 Septomaxillae_2 / 'present, contacting each other at midline, the maxillae anteriorly and lying on the top of the vomers' present_as_small_slivers_of_bone_lying_within_the_nasal_cavity_that_do_not_contact_each_other_at_midline absent, 42 Nasal_median_contact_3 / separated_by_less_than_50%_of_their_length separated_for_more_than_50%_of_their_length_by_nasal_process_of_premaxilla completely_separated, 43 Posterior_margin_of_nasals_4 / 'posterior aspect of nasals form a smooth curve, with apex in the middle of the bone' posterior_aspect_of_nasals_pointed posterior_aspect_of_nasals_squared, 44 'Prefrontal-nasal relationship 5' / broad_contact_between_prefrontals_and_nasals 'small area of contact between prefrontals and nasals, with nasals separated anteriorly from prefrontals by prefrontal process of maxilla' 'prefrontals completely separated from nasals by prefrontal process of maxilla and antero-lateral process of frontal bone', 45 Anterolateral_spine_of_prefrontal_6 / absent 'present, projecting anteriorly over the premaxillary process of maxilla', 46 'Groove at prefrontal-lacrimal junction 7' / absent present, 47 'Palatine-ectopterygoid relationship 8' / no_contact_between_ectopterygoids_and_palatines 'ectopterygoids contact palatines; excluding maxillae from infraorbital foramen', 48 Palatine_process_of_pterygoid_9 / does_not_project_past_the_anteromost_border_of_infraorbital_foramen extends_anteriorly_beyond_the_infraorbital_foramen, 49 Quadrate_process_of_pterygoid_10 / 'tapering posterioly, of with parallel ventral and dorsal edges, but not expanded' quadrate_process_of_pterygoid_expanded_terminally, 50 'Shape of mid-sagittal section of frontal 11' / anterior_and_posterior_aspects_of_frontal_bone_curve_ventrally 'frontal bone is flat in mid-sagittal section view, or anterior and posterior aspects of frontal bones curve dorsally', 51 Squamosal_process_of_julgal_13 / thin_squamosal_process broad_squamosal_process, 52 Posterior_angle_of_jugal_15 / 'not expanded postero-ventrally, rounded' 'expanded postero-ventrally, with postero-ventral aspect of jugal approaching the posterodorsal aspect', 53 Supraorbital_process_16 / 'absent; postorbital and prefrontals lack processes' 'intermediate condition; prefrontals have posteriorly directed process and postorbital have anteriorly directed processes' postorbitals_in_contact_with_prefrontals_via_a_supraorbital_arch, 54 Lateral_shelves_of_adductor_crest_18 / absent_or_when_present_is_small_and_restricted_to_the_anterolateral_aspect_of_the_adductor_ridge present_along_the_lateral_aspect_of_the_parietal_crest 'large, expanded laterally to partially overlie the supratemporal opening', 55 Development_of_the_parietal_blade_20 / 'post-embryonic development of median parietal blade' embryonic_development_of_parietal_blade, 56 Parietal_blade_sexual_dimorphism_21 / 'parietal blade well-developed in males only' 'parietal blade well-developed in both sexes', 57 Expansion_of_parietal_blade_22 / absence_of_a_parietal_blade dorsal_expansion ventral_expansion, 58 Dorsal_process_of_squamosal_23 / 'prominent dorsal "hook" contacting the supratemporal' 'no dorsal "hook", i.e. ventral projection equals dorsal projection', 59 Squamosal_24 / 'curves dorsally, not contacting or overlapping the anterodorsal aspect of the quadrate (single articulation)' 'straight, overlapping the anterodorsal border of lateral concha of quadrate, with sharp laterally pointed spine (double articulation)', 60 Medial_concha_of_quadrate_25 / 'expanded medially past the cephalic condyle; the medial concha may or may not contact the antero-lateral process of the paraoccipital process' 'not expanded, past the condyles (constricted)', 61 Quadrate_26 / posterior_crest_arches_anteriorly posterior_crest_is_vertical_with_no_anterior_curvature, 62 Epipterygoids_27 / contacting_the_ventral_portion_of_parietal free_or_contacting_the_alar_process_of_prootic, 63 Crista_ventrolateralis_28 / 'broad, expanded laterally obscuring the vidian canal' 'narrow, not expanded laterally', 64 Outlines_of_bony_labyrinth_29 / 'superficial outlines of bony labyrinth obscure, not raised above the surface of the occiptal bones' intermediate_condition 'superficial outlines of bony labyrinth very distinct,  raised well above the surface of the occiptal bones', 65 Posttemporal_fenestra_30 / 'open, with the squamosal process at an angle of more than 90 degrees' partially_closed 'closed, owing to the limiting angle of the squamosal process of parietal (less than 90 degree)', 66 Coronoid_lateral_process_33 / 'coronoid bone with a small, irregular process overlapping the posterolateral surface of dentary' a_posteriorly_directed_curving_process_descends_over_the_posterodorsal_surface_of_the_dentary 'coronoid bone without a process overlapping the postero-lateral surface of the dentary', 67 'Size of posterior aspect of angular (labial view) 34' / extends_between_coronoid_process_and_articular_fossa 'lateral aspect reduced, does not extend between the coronoid process and the articular fossa', 68 Angular_process_size_35 / 'well defined, in juveniles and adults (non-transforming ontogeny)' 'small in juveniles and well-defined only in large specimens', 69 Angular_process_position_36 / angular_process_positioned_in_a_horizontal_plane positioned_in_a_oblique_plane, 70 Hyoid_apparatus_pattern_37 / 'z-pattern in wich the second ceratobranchial is the shortest posterior element, the first ceratobranchial and the ceratohyal being of equal length' 'x-pattern; the second ceratobranchial is the longest element, the first ceratobranchial and the ceratohyal may or may not be equal in length', 71 Rostral_position_of_skull_38 / arched_dorsally 'rostral position of skull is pointed or keel-shaped (spatulate)', 72 Crowns_of_posterior_marginal_teeth_39 / 'tricuspid, with more or less paralell sides' 'tricuspid, with distinctly expanded crowns' 'tricuspid, with distinctly tapered crowns and very small secondary cusps', 73 Number_of_presacral_vertebrae_40 / 24. 23. 22., 74 Lumbar_ribs_41 / all_with_free_articulating_ribs one_or_more_with_ribs_absent, 75 Ratio_of_number_of_neural_spines_to_number_of_transverse_process_in_caudal_verte / '1$1 ratio' '2$1 ratio', 76 Neural_spine_height_44 / neural_spines_same_size_as_vertebral_body 'neural spines enlarged in thoracic, lumbar and caudal regions in males' 'neural spines reduced, shorter than height of vertebral body', 77 Rib_articulation_patterns_45 / '6(4+2)' '7(4+3)' '5(3+2)' '6(3+3)', 78 Suprascapular_cartilages_46 / 'project dorsally to the level of the vertebrae, sopmetimes meet above neural spines' suprascapular_cartilages_do_not_reach_the_level_of_the_vertebral_column, 79 Dorsal_extent_of_clavicle_47 / 'clavicle reaches the suprascapular cartilages, with the acromion process located on suprascapular cartilage or at the scapulo-suprascapular junction' 'clavicle does not reache the suprascapular cartilages, with acromion process located on anterior aspect of scapula', 80 Clavicular_lateral_margin_48 / 'lateral margin of clavicle irregular, or forming a smooth curve' 'lateral margin of clavicle distinctly angular, with an acute projection at the apex of the angle', 81 Clavicular_fenestrae_49 / absent present, 82 Preacetabular_process_50 / prominent reduced, 83 Anterior_extension_of_symphysis_pubis_51 / symphysis_pubis_does_not_extend_anteriorly_past_the_pubic_tubercles symphysis_pubis_extends_anteriorly_past_the_pubic_tubercles_with_pubic_rami_join_together_forming_an_acute_angle, 84 Infratemporal_fascia_52 / 'fascia present, variably covering the posterodorsal aspect of the infratemporal fossa to the entire M. adductor mandibulae externus' little_fascia_present_at_posterodorsal_corner_of_infratemporal_fossa, 85 Insertion_of_M._adductor_mandibulae_externus_53 / mandibular_insertion_projects_past_the_surangular mandibular_insertion_restricted_to_surangular, 86 M._levator_anguli_oris_54 / origin_includes_the_tympanic_crest_of_the_quadrate does_not_originate_from_the_tympanic_crets_of_the_quadrate, 87 M._adductor_mandibulae_externus_superficialis_55 / single_dorsal_head two_separate_dorsal_heads, 88 M._adductor_mandibulae_externus_profundus_56 / dorsal_head_well_developed dorsal_head_reduced dorsal_head_is_absent, 	89 Superciliary_scales_57 / elongate_and_strongly_overlapping intermediate 'quadrangular, nonoverlapping'
	;
	MATRIX
		Corytophanes_hernandezi         00113101011000010021101001010010001010011111100101111211211111021111101001002001000011021
		Corytophanes_percarinatus       0011310101000001002110100101001000101001(12)1111001011122112111110220111110(01)1000011000011021
		Corytophanes_cristatus          0011310101?000010021101001010010001010011212101101112211210111022011111001000111000011021
		Laemanctus_serratus             01103?00000000000021101001010010001010001020010011010111210110020101101001023000100100112
		Laemanctus_longipes             01103?00000000000021101001010010001010001000010011010111210110020101101001021000100100112
		Basiliscus_plumifrons           00012100000000000021101000000010001010100000010010000000100000110000010100110000111000000
		Basiliscus_galeritus            000121000000000000211010000000100010101000000100100000001000001100000?0100100000111000000
		Basiliscus_vittatus             00012100000000000021101000000010001010000000010010000000100000110000000100100000111000000
		Basiliscus_basiliscus           00012100000000000021101000000010001010000000010010000000100000110000000100110000111000000
		Polychrus_acutirostris          110000011100111101??10221101?1000112000010000000000000000101000201101110000010100(01)(01)(01)00002
		Tropidurus_etheridgei           0000000002112000010001110010011010020100(01)01000000000000001000000021(01)10010000100(01)0(01)0010(01)00
		Sauromalus_obesus               000010100100000000100020?00100010000001000(012)00000000000000000000(01)020(01)1(01)010000000101001000(01)
		Elgaria_panamintina             000000000?0?00?010???0?0000?1000000?000??????????????????????????????????????????????????
	;
END;

BEGIN ASSUMPTIONS;
CHARSET FrostEtheridge = 1 - 39 ; 
 CHARSET Lang = 40 - 89 ; 
 CHARSET Total Gross-Morphology = 1 - 89 ; 
 OPTIONS DEFTYPE = unord PolyTcount = MINSTEPS ; 
 TYPESET * UNTITLED = unord: 1 - 6 8 - 9 11 - 12 14 - 21 26 - 35 37 - 40 43 45 - 53 55 - 63 67 - 72 74 - 75 77 - 87 , ord: 7 10 13 22 - 25 36 41 - 42 44 54 64 - 66 73 76 88 - 89 ; 
 WTSET * UNTITLED = 999: 1 - 89 ; 
 END;