#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on June 13, 2026; 15:38 GMT TreeBASE (cc) 1994-2008 Study reference: Koopman W., Wissemann V., Cock K., Huylenbroeck J., Riek J., Sabatino G., Visser D., Vosman B., Ritz C., Maes B., Werlemark G., Nybom H., Debener T., Linde M., & Smulders M. 2008. AFLP Markers as a Tool to Reconstruct Complex Relationships: A Case Study in Rosa (Rosaceae). American Journal of Botany, 95(3): 353-366. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S2030] [ The following blocks are input data for analysis step 4210 ] BEGIN TAXA; TITLE TaxaForAnalysisStep4210; DIMENSIONS NTAX=128; TAXLABELS Rosa_Aimee_Vibert_416 Rosa_Alba_Suaveolens_407 Rosa_Aloha_424 Rosa_Arthur_de_Sansal_419 Rosa_Blush_Damask_405 Rosa_Catherine_Mermet_415 Rosa_Chateau_de_Namur_402 Rosa_Climbing_Cecile_Brunner_427 Rosa_Communis_401 Rosa_Diablotin_428 Rosa_Felicite_Parmentier_406 Rosa_Fellenberg_417 Rosa_Guirlande_dAmour_434 Rosa_James_Mitchel_410 Rosa_Kathleen_Harrop_412 Rosa_La_Reine_421 Rosa_La_Ville_de_Bruxelles_404 Rosa_Le_Rire_Niais_408 Rosa_Little_Buckaroo_431 Rosa_Mignonette_426 Rosa_Miranda_418 Rosa_Nebada_432 Rosa_Nozomi_430 Rosa_Pink_Chimo_438 Rosa_Prosperity_435 Rosa_Queen_of_Bourbons_413 Rosa_Rose_de_Meaux_409 Rosa_Scharlachglut_439 Rosa_Slaters_Crimson_China_422 Rosa_Superb_Tuscan_403 Rosa_Triomphe_de_Luxembourg_414 Rosa_Violette_437 Rosa_Yolande_dAragon_420 Rosa_abietina_202 Rosa_acicularis_106 Rosa_acicularis_67 Rosa_agrestis_146 Rosa_agrestis_160 Rosa_agrestis_203 Rosa_arkansana_445 Rosa_arvensis_18 Rosa_arvensis_2 Rosa_arvensis_204 Rosa_blanda_447 Rosa_caesia_205 Rosa_caesia_44 Rosa_canina_147 Rosa_canina_156 Rosa_canina_4 Rosa_canina_69 Rosa_canina_Pfander_71 Rosa_canina_Schmids_Ideal_72 Rosa_canina_Superbe_70 Rosa_canina_var._andegavensis_206 Rosa_centifolia_Blanche_Moreau_101 Rosa_corymbifera_17 Rosa_corymbifera_207 Rosa_corymbifera_32 Rosa_corymbifera_Laxa_201 Rosa_corymbifera_Laxa_36 Rosa_corymbifera_Laxa_76 Rosa_damascena_Ispahan_10 Rosa_dumalis_100 Rosa_dumalis_148 Rosa_dumalis_208 Rosa_elliptica_209 Rosa_foetida_30 Rosa_gallica_210 Rosa_gallica_Boule_de_Nanteuil_7 Rosa_glauca_163 Rosa_glauca_211 Rosa_henkeri_schulzei_162 Rosa_hugonis_440 Rosa_inodora_212 Rosa_jundzillii_213 Rosa_majalis_164 Rosa_majalis_214 Rosa_majalis_40 Rosa_majalis_Fulgens_39 Rosa_micrantha_151 Rosa_micrantha_215 Rosa_micrantha_216 Rosa_mollis_217 Rosa_moschata_436 Rosa_multiflora_55 Rosa_multiflora_58 Rosa_multiflora_Inermis_53 Rosa_nitida_442 Rosa_pendulina_218 Rosa_persica_451 Rosa_pseudoscabriuscula_152 Rosa_pseudoscabriuscula_219 Rosa_roxburghii_444 Rosa_rubiginosa_220 Rosa_rugosa_49 Rosa_sericea_var._omeiensis_16 Rosa_sertata_96 Rosa_sherardii_165 Rosa_sherardii_166 Rosa_sherardii_221 Rosa_spinosissima_222 Rosa_spinosissima_93 Rosa_spinosissima_Kakwa_29 Rosa_stylosa_223 Rosa_subcanina_153 Rosa_subcanina_224 Rosa_subcollina_155 Rosa_subcollina_161 Rosa_subcollina_225 Rosa_sweginzowii_Macrocarpa_63 Rosa_tomentella_154 Rosa_tomentella_226 Rosa_tomentella_78 Rosa_tomentosa_227 Rosa_turcica_77 Rosa_villosa_228 Rosa_villosa_23 Rosa_villosa_mollis_24 Rosa_virginiana_443 Rosa_wichuraiana_Temple_Bells_61 Rosa_wichurana_59 Rosa_wichurana_60 Rosa_willmottiae_Wisley_Farm_43 Rosa_woodsii_50 Rosa_woodsii_92 Rubus_caesius Rubus_fruticosus Rubus_phoenicolasius ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M3135] TITLE AFLP_Fig._1; LINK TAXA = TaxaForAnalysisStep4210; DIMENSIONS NCHAR=520; FORMAT DATATYPE=Standard SYMBOLS= "1 0" MISSING=? GAP= -; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 200 210 220 230 240 250 260 270 280 290 300 310 320 330 340 350 360 370 380 390 400 410 420 430 440 450 460 470 480 490 500 510 520] [ . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .] Rosa_Aimee_Vibert_416 1000100000000010000100000100011000000000000010110100001000000100000000000000000100100000000000000000000100000100000000100000000000000110001000000000001101001001010000000110000000000000000000000000000000000000000000000000000000100101000100000010000010001000000000001000001010000011110010100000000001100000100100010000111000011010000000000011011110111000111011111010001011000110101011110010010001111110101110000001100111100100100111101100001100010010100000101000001111010010101011000001011010101010011011000000111001101111 Rosa_Alba_Suaveolens_407 1000100000000010010000000100010000000000000000010000001000000100001000000000001100000000010000001000000100000100100000100010000000001010001000000000000111101011011000100110000101100000001010001000000110000000011000000000000000100100111000000010100010010000000010100000001010000001000010000000100001100000100000000110111000011110100100000010?11111011100111011111011111011010101111111110110101111111100111110011001001011?01101111110011011111111111110100111111011101111010100110101101111100011001010001011010000111101100111 Rosa_Aloha_424 10000101001001100111000000000011000000100000000111000011000001000010100000000011001000000000000010000011000000000000001000000000000000000000100010011010010011010110000000100001000010000000000010000000100000000100000000010000001001001000000000101000100010100000001010000010100000011010101000000000000000001000000000001110000111100000000000111110101001001111111110100011110011011001100101100110011110101100101000101101100010000001001010010111010111001001111011010110110110010000100010110?0010100011111101010000001001101111 Rosa_Arthur_de_Sansal_419 0000100000000010000100000100000000000000000000010000011000000110001000000000000100000000000000000000000100000100100000100010000000001010001000000000000101100001010100000010000000001000000010001000000010000000000001000000000000100100000000000010000010000000000010001000001010000001000010000000100001000000100000000000111010011111000000000000001010000101111011101011001011100101111111110110100111111100111100000000000011011100000100100100011110011000100011101000011110111100110110001111010001101011011011010000111001101110 Rosa_Blush_Damask_405 1000100000000000000110000100100100000000000000010000001010000010001000000000000100000000000001000000000100000100000000100010000000001011001100000000000101100000011000000110000100101000000010001000000100000000000100000000000000100100011000000010000010000000000010101000000010000001000010000000000001000000100000000000111010011011100100000010011110000101111011100011101011010001111101110110100111111100101110000000110010100101110111001110100011011110100101101111101111011010110111001111101111110000011011010000111101101111 Rosa_Catherine_Mermet_415 1000100000000000000100000000001100000000000000110000011100000100000010000000001100000000000000000000000000000100000000100000000000000010011000001001001001001001010000000010000000010000000000001000010010000101000001000000000000000100000000000010000010001000000000001000001010100001000010100000100000000000101000000010111000011110000100000011001010010000110111111110101011011011111111110010111001111000110001000000110110110101011110101100011111011110100011101100011101111010000011000110110110100011010111010000101101101111 Rosa_Chateau_de_Namur_402 10001001000000100000000001000100000000000001000111000010000001100011000000100001000000010000010010000001010000001000001000100000000010110010100010000000011000000101100000100110111111001000100010011001100000010111010000000010001001001010000000100000100000000000011000000010100000001000111000001000010000001000000010001110100111111001000000111101110111011111111000111110010110111111011100?011011011011001110100000010110100000110000001111011?110111111100111111001011000111110101010011001110011000011001001010000111101101110 Rosa_Climbing_Cecile_Brunner_427 0000110100000010001000000000000100000001000010011000001100110100000000000000000110000000000000001000000001000100000000100000000000000000000010001000000001000000010000000000000000000000100000001010100010000000000000000000000000000100000000000010000010011000000000100000001010000000000010000000100001000000101000000000111000011010000000000010111011110011101001100100110011?011001101100101100111111010101110110001110101001010010011111010011000100110000001001111000110111111100111110110111110101110101??????????????????????? 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Rosa_Fellenberg_417 100001011011001000110000000000010000000000000001100000110010010000001000000000011010000000000000000000000000010000000010000010000000001000001000001010100100000101001000001000010000010010000000101000001010000100100000000100000000010010010000101000001000100000001010100000101010000100001000000010000000000010100000000011100001111000000000001001001010000011101111011011100100100011110001111101111111101111111100000000000000000000010000000001010000000??01111111010101001111011101111011111101010101001110100010000001101101111 Rosa_Guirlande_dAmour_434 100010000000000000010000000001110000000100001001100010100000010000000010000010000000000000000000000000000000000000110010000000000000000001100000000000100110000100000000001000000010000000000000100000000000010000000000000000001010010000000000001000001001001000000000100000101000000101001010000010000100000010000000000001100001111000000000000111101100111011001111101011100110?111101010111111101011100110101111000000100101001011000100001000001001111011?01011101000001001001010110010010100100001100010010101010000001001100110 Rosa_James_Mitchel_410 1000100000000010000100000100010000000000000000010000011000000110001000000000001100000000000001000000000100000000100000100010000000001011001000000000000001101001010000000110000101101000000010001000000110000000000100000000000000100100001000000010100010000000000011000000001010000001000010000000100001000000100000000010111010011111000100000010011010000101111001111011101011010101111111110110110111111100101100000000100011010101010110011010100101011010101001101001101100011110111010001111110001101011011011010000111101101111 Rosa_Kathleen_Harrop_412 1000010100110010010000000000001000000000000110111100001000110100000000000000000100100000000000001000000000010100000000100000000000000001000010111000011101001000001010000010001010001100101000001010100010010101000000000001000000100100100100000010100010001000000000100000001010000000101010100000000000000000100000001010111010011110000000000011?11110110100111001111110111011001011111110010110011101101000110000100000001100100011000101101001?00001001100100001111001011000101000100010000010010100110011001101010000111101101111 Rosa_La_Reine_421 1000010000000000000100000100001000000000100000110100011100100100101000000000001100100000000000000000000000000100000000100000000000001010000000011000101101100001010000000010000000000000000010001000000010000000000000000001000000100100001000000010000010000000000010001000001000000001000010000000100000000000100000000000111010011111000000000011011010000000111011111011001011101001111110010110100011111101111000000000100111011101000100101100011111111100100010011000011001110001000110001011110000110011111011010000111001101111 Rosa_La_Ville_de_Bruxelles_404 1000100000000010000100000100101000000000110100010000001010000110001000000000000100000001000001000000000100000000000000100000000000001011001000000000000101100001011000000110000101101000000010001000000110000000010100000000000000100100000000000010000010000000000011001000001010000001000010000000100001100000100000010010111010011111000000000010?1111011110111101110001110101101110111111011111010000111110010101000100011001000010111011100111010101111111010000?100000101101011010101010001111110011010010011011010000111011101110 Rosa_Le_Rire_Niais_408 100010000000001000010000010001000000000000010001000000100000010000100000000000110000000000000100000000000000010010000010001000000000101000110000000000000110000101000000011000010010100000001000100000011000000001010000000000000010010000100000001010001000000000001000000000101000000100001000000010000110000010010000001011101001111110010000001001111000010111001111101110101101010111111111?110110011111110101010000001001001001101110111011111101111111110100011100001101011011100111110000101110011110011011011010000111011101111 Rosa_Little_Buckaroo_431 1100100000000000000100000100001100000010000000110010001110000100000010000000000100100000000000000000000001000000000010100000000000000010001100000000001001000001000000000010001000010000000000001000000000000000000000000000000000100100000000000000100010000001000000001000001010100001000110000000000001000000101000000000111000011110000000000011011100111010111010111001101111101011101110010010010001110100011010000000100101001101011110101101011111110110101000101000001101110111101010010111010010100011010110010000011001100111 Rosa_Mignonette_426 10001000000000000001000000000011000000000001001110000010100001000000000000000001000000000000000000000000000000000011001000000000000000000110000000000010011000010100000000100000000100000000000000000000000001000000000000000000101001000000000010100000100000000000000010000010100000010001101000001000011000001010000010001110000111100000000000111111000010101101111110101010011010101010100100101000011101000011000000000001000010000000000001010111000100110000001010100110111101011000100101001000110000100??????????????????????? 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Rosa_subcollina_155 00001000000010000100000001000000000000000000000100000010100000110010000000000101000000000100010000000011100001000000001000110000100000100110000000000000011000010101001000100000000000000000000100000001000000000010000000000000001001000110000100101000100000000000000001000010100000000000101000000000010100001000001010101110000111100000000001100000?0110101011110111011110111010100110000100010000011010100101110010100010100000110111000001?10100111010100?00000001000001100001000110011100101?11110001000000111110010111111101111 Rosa_subcollina_161 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10001000000010000100000001000000000000000000001000000110100000111010000000000101000000000100010000000001100001001000001000110000100000100010000000000001011000010111001000100000100000000000000000000001000000000010000000000000001011000110000000101000100000001100000001000010100000000000100000000000010000001000000010101110000111100000000001101010101?010001101011101011011100111011010011001010010101?101101110010000000000000111111000011110100110010100?11000111000111111011010110111100101111010001000000111100010111101101111 Rosa_sweginzowii_Macrocarpa_63 00001000000000000100000001000001000000000000101101000010100000000000000000000000100000000100000000000000000000001000011000100000001000000010000000001000010000100100000000000000000000000000000010000001000000010000000000000000001010000110000010011000000000000000000001000011100000000000100000100000010000001000000010101101000111100000000000000100110010010111001100??101011??00000?01001?0111110001000100001001000000000000001100011100001101000000000101000000000001000000100011100111000001101010000000110101000010011001101111 Rosa_tomentella_154 0000100000001000000000000100000000000000001000010000001010000111001000000000010100000001010001000000010010000100000000100010000010000010001000000000000001100000011100100010000000000000000000000000000100000000001000000000000000100100011000010010100011000000000000000100000010000000?00010100000000001000000100100101010111000011110110000000110000000100100011000111001110111010100110100000010100001111101101110010000000011000010010000001110100110010100?00000000000000000001000100100101000001100001000000111100010111101101111 Rosa_tomentella_226 100010000000101001000000010000000000000000100010000000101000001100100000000001010000000001000100000000011000010000000010001100001000001100100000000000010110100101010010001000000000000000000000000100010000000000100000000000000010010001100001001010001100000000000000010000001000000000001000000000000100000010000110101011100001111000000000011000101011000?011100111010110111011110110100110110100101111101111110011100000001000111110000001000111110000100?0000011100000111101101011011?100101011010001000000101100010111101101111 Rosa_tomentella_78 1000100000000000000000000100000000000000000100010000001000000010001000100010010100000000000001100100100100000100000000100000000000000011001000000000000001100000001000000010000001001010000000001100000010000010010100000000000000100100000000000010100010001000000000001001001000000000000010000000000000000000100000101000111010011111100000000010011110100101111001100000010?11000011100100011110000111100100111000000000000011000101100100001010100011010001100011100000111011011000100110000000011011110010011011010010111001101111 Rosa_tomentosa_227 100010000000100001000000010000000000000000000000000000101000001100100000000001010000000001000100000000010000010010000010001000001000001000100000000000000110100101000010001000000000000000000000000100010000000001100000010000000010110001100001001010001000100000000000010000101000000000001000000000000100000010000010101011100001111000000000001001101011000?0110101110101111110011101001001101101001010??101101110011110000011000111111000011111101011010110?0100011100?101111111010111111100001111011001000010111110010111101101111 Rosa_turcica_77 000010000000000000000000000000000000000000000001000000100000001000100000001001010000000000000100000000010000010000110010000000000100001000100000000000010110000000000000001000000100100000001000100000001000000001010000000000000010110000000000001000001000100000000000100100001000100000001000000000000000000010000000100011101001101110000000001000011000000?111010100000000011000011100000010110000011100100101000000010000000000101110100001010100011000000100010101000001011011000101110000000010011100010011011010010111001101111 Rosa_villosa_228 100010000000000001000000010000000000000000000001000000101000000100100000000000010000000001000100000000010000010010000010001000001000001001100000000000010110100101000010001000000000000000000000000000010000000000100000000000000000010001100010001010001000000010000000010000101000000000000000000000001100000010000000101011100001111000000000000001000000??00011100101010000000000100100100100110000111011101101110011010010100000101111000011100100010010010?1111111101110?111111011101111101001011010011000010111110010111101101111 Rosa_villosa_23 10001000000000000100000001000000000000000000000100000010100000010010000000000101000000000100011000000001000001001000001000100100100000100010000000000000011000010100001000100001000000000000010100000000000000000010000000000000000001000110001000101000100010001000000001000000100000000000000000000000010000001000000010101010000111100000110100101110101001010111101111111010101010100?01011111100001110111011010100100100001100001011101000111011000101100?0?000010100100001110010101?011110000101101?001000010101100000111101101110 Rosa_villosa_mollis_24 00001000000000000100000000000000000000000010000000000010100000010010000000000101000000000100010000000001000001001000001000100000100000100000000000000000010000000110000000000000000000000000000000000001000000000010000000000000000001000110000000100000100000000000000001000000100000000000100000000000000000000000000010101010000111100000000000100000001001000100101110111110101010001000001101100001110111001010100110000000000001001000000110011000100100?0?00000000001000111001000000111100101011000000000000001100000111101100111 Rosa_virginiana_443 100000100000010001010000000000010001100000000001000000101100001000000000000000000000000001000000000010010000000000010010001000010000000000011000000000000110010101100000000000000010000000000000100000010000000000000000000000000010000011110000000010100000100000000000010000100000000100001000000000000010000010000000001011000001101000000000000000011110000111111111100111111110110010010001001000000110111010011000000000000100111100010100101001010111111??00101000101000101010111101110100110001111011111010101010000100101100111 Rosa_wichuraiana_Temple_Bells_61 10001000000000000000000101000001000000100000000110100010000001000000000000000001000000000000001000000010000000000100001000000000000000000010000000000001011000000100100010100000000000000000000000000000000000000000000000000000101011010000000010100000100000000000000010000010100000000001100000000000010000001000001010001110000111100000000000011111000111000010100101011?1010101001101011110110?00111110110001101000000000100010101001110001000001100010010000001000000011010111011000010010011110010100010010101000010001001101110 Rosa_wichurana_59 1000100000000000000000010100000100000010000010001000001000000100000000000000000100000000000000100000001000000000010010100000000000000000011000101000000101101000010000000010000000000000000000000000000000000110000000000000000010101100000000000010000010000000000000001000101010000000000110000000000001000000100000001000111000011110000000000001111100?0110?0100100?00011?0010101001101???????????????????????????000000000100010101000?1000100000110001000000000000000001111011111?100010011000100010100000010101000010001001101110 Rosa_wichurana_60 1000100000000000000000010100000100000010000100011000001000000100000000000000100100000100000000000000001000000000010010100000000000000000001000000100000001101010000000001010000000000000000000000000000000000100000000000000000010100100000000000110000010000000000000001000001010000000000110000000000001000000000000001000111001111110000000000001111100?01100011?100101011?0110101001101001010010?00111110110001101000000000100000001000?10001000001100010000000000000000001000101011100010000001100010100010010101000010101001101111 Rosa_willmottiae_Wisley_Farm_43 1000001000000000000000000100000100000000000010000000001010000000000000000000000000000000010000000000010100000000000000100010000000000000000010000000000000000000011000000000000000010000000000000000000100001000010000000000000000000000011000000000100000000000000000000100001000000000010000000000000000100000100000001010110101011010000000000000010000110000111000111010011110000000110101100010011111111110000000110010000010000111111100001111001110010100100000000010010100010000100000000101000000000000010101000000101101101110 Rosa_woodsii_50 100000100000000000000000010000000001100000000001000000101000011000000000000000000000000001000000000000010000000000000010001001000000000000001000000000000100000001100000000000001000000000000000000000010000011000000000000010100010000101100010000010000000100000000000010000100000000000100000010010000100000000010000100011000001111000000000001001000100010011101110011011101110110010?100110010110111100110000100000000010011100100000110001111000100011010010011001111000111010011?10001000101011011011100101001100000101100100111 Rosa_woodsii_92 10000010001000000100000001000001000000000000000000000010100000100000000000000000000000000000000000000001000000000001001000100000000000000000100000000000011000000100000000000000000000000000000000000001000000000000000000000000100000010110001100001000000000000000000001000010000000000000000010000000100000001000001010001100000110100000000000000000010011101011?11000001000111001000001000100100001010101101001011000000000000001000010000011110000000100000000010000000100??000010110011000101001100001010010011000000110101100111 Rubus_caesius 1010100000000001100001000001000001000100001000?00000?000100001000000?0010001000000001000100000000001100000000?1000000010000000000000010001100100001000100000000000000100000101000000100000010000000000101001000001000100101001000100000000001000010001000000010100100000000011010000100001011100000100100000000000100000010100?000100010000000100001100011001100000??010100101101101?001001001?11001?01100011100?010?1000001000010000000010100000011000000101100110000000010000000001111?01010101??001011000?10?1000001100?1010000010?11 Rubus_fruticosus 10001000000000000000000001101001000000000000?0000000?000000001000001?01000000000010000000000000000000000001000000000001000000010000000000010000001000001000101001000010100100000000000000000000000100000000000000100001100000000000000000110010000001011000000000010000010100000000000000000001000?00?01000000000001000001010010100001100000001000010101010001010000001010011?101101?100001001111000?1100?1111111000000000110010001000000001000000001010001111011001000100100000000001100001001111010000100101011000000010?0000000000010 Rubus_phoenicolasius 0000100000000000000001000000000000000000000000000000000000000100000000000000000000001000000011000000100000000110000000000010?000000000001010011000000000000100000100010001?00000000000000000000000000000000000000000000000000001000000001000000000000000000001000000000110000000000000000000100000000?000000?0000000000110001100000000100010000000000000?00000000001001011000010?101?01100010001100010100000111010110000000000000000000100000000010100100000000010000000100000000000000?000000000000000000?0?0000000000000?0000000001010 ; END; [ The following blocks are output data for analysis step 4210 ] BEGIN TREES; TITLE Tb8982; LINK TAXA = TaxaForAnalysisStep4210; TREE Fig._1 = [&R] ((((((((((((((((((((((Rosa_canina_var._andegavensis_206,Rosa_corymbifera_207),Rosa_caesia_205),(Rosa_subcollina_225,Rosa_tomentella_226),Rosa_subcanina_224),Rosa_dumalis_100),(Rosa_corymbifera_Laxa_201,(Rosa_sherardii_165,(Rosa_abietina_202,(Rosa_sherardii_221,(Rosa_tomentosa_227,Rosa_pseudoscabriuscula_219))))),Rosa_dumalis_208),(Rosa_subcollina_155,(Rosa_canina_156,(Rosa_dumalis_148,Rosa_subcanina_153)))),Rosa_jundzillii_213,(Rosa_canina_147,Rosa_agrestis_146),(Rosa_subcollina_161,Rosa_sherardii_166)),(Rosa_agrestis_160,(((Rosa_agrestis_203,Rosa_inodora_212),Rosa_elliptica_209),(Rosa_henkeri_schulzei_162,(Rosa_micrantha_215,(Rosa_micrantha_216,Rosa_rubiginosa_220)))))),((Rosa_corymbifera_32,Rosa_caesia_44),(Rosa_corymbifera_17,(Rosa_canina_4,Rosa_corymbifera_Laxa_36)))),Rosa_stylosa_223,((Rosa_canina_69,Rosa_canina_Pfander_71),Rosa_canina_Superbe_70)),((Rosa_villosa_23,Rosa_mollis_217),Rosa_villosa_228)),(Rosa_micrantha_151,Rosa_pseudoscabriuscula_152,Rosa_tomentella_154)),(((Rosa_canina_Schmids_Ideal_72,Rosa_corymbifera_Laxa_76),Rosa_glauca_211),Rosa_glauca_163)),Rosa_villosa_mollis_24),((((Rosa_arvensis_2,Rosa_arvensis_204),Rosa_arvensis_18),((Rosa_turcica_77,Rosa_tomentella_78),Rosa_gallica_210)),((Rosa_Chateau_de_Namur_402,Rosa_Scharlachglut_439),Rosa_Felicite_Parmentier_406,((Rosa_gallica_Boule_de_Nanteuil_7,Rosa_damascena_Ispahan_10),Rosa_centifolia_Blanche_Moreau_101,(((Rosa_Miranda_418,Rosa_Arthur_de_Sansal_419),(Rosa_Yolande_dAragon_420,Rosa_La_Reine_421)),(((Rosa_Superb_Tuscan_403,Rosa_Blush_Damask_405),Rosa_La_Ville_de_Bruxelles_404),(Rosa_Alba_Suaveolens_407,(Rosa_Communis_401,(Rosa_James_Mitchel_410,(Rosa_Le_Rire_Niais_408,Rosa_Rose_de_Meaux_409)))))))))),((((Rosa_wichurana_59,Rosa_wichuraiana_Temple_Bells_61),Rosa_wichurana_60),(Rosa_multiflora_Inermis_53,Rosa_multiflora_55)),(((((((Rosa_Aloha_424,Rosa_Diablotin_428),Rosa_Slaters_Crimson_China_422),(Rosa_Kathleen_Harrop_412,Rosa_Queen_of_Bourbons_413)),Rosa_Nebada_432),Rosa_moschata_436),(Rosa_Fellenberg_417,Rosa_Climbing_Cecile_Brunner_427)),(Rosa_Nozomi_430,((((Rosa_Triomphe_de_Luxembourg_414,Rosa_Catherine_Mermet_415),Rosa_Little_Buckaroo_431),Rosa_Aimee_Vibert_416),((Rosa_Pink_Chimo_438,Rosa_Prosperity_435),(Rosa_Guirlande_dAmour_434,(Rosa_Mignonette_426,Rosa_Violette_437)))))))),((Rosa_spinosissima_Kakwa_29,Rosa_spinosissima_93),Rosa_spinosissima_222),(((Rosa_nitida_442,Rosa_virginiana_443),(Rosa_arkansana_445,Rosa_blanda_447)),(Rosa_sweginzowii_Macrocarpa_63,(Rosa_willmottiae_Wisley_Farm_43,((((Rosa_majalis_214,Rosa_majalis_164),Rosa_majalis_40),(Rosa_majalis_Fulgens_39,Rosa_rugosa_49)),(Rosa_woodsii_92,((Rosa_pendulina_218,Rosa_sertata_96),(Rosa_acicularis_106,(Rosa_woodsii_50,Rosa_acicularis_67))))))))),(Rosa_hugonis_440,Rosa_roxburghii_444)),(Rosa_persica_451,(Rosa_sericea_var._omeiensis_16,Rosa_foetida_30))),Rosa_multiflora_58),(Rubus_fruticosus,Rubus_caesius)),Rubus_phoenicolasius); [! TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr3017] END;